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Internet Directory of Biospeleologists
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Directory of e-mail addresses and/or Home Pages of biospeleology researchers.

Because this is a field where they have made and continue to make valuable contributions, amateurs (in the proper sense of the word), are included as well as professional investigators affiliated with research institutions. I would like to make this directory as comprehensive as possible, and invite anyone working on any aspect of biospeleology to send their details to

If using the email links here, note that the address in your mail program will have to be edited. Simply remove the "*AT*" and replace with "@". This is a simple anti-spam measure which may be effective.


  • Aulenbach, Nancy H. (Alabama, USA). email . Biological inventory of a new cave in Alabama.

  • Biswas, Dr. Jayant (India). email . Cave biology in India. web page: Cave Biology webpages

  • Borowsky, Dr.Richard L. (New York, USA). email . Genetics of cave fishes. Affiliation: New York State University.

  • Boston, Penelope J. (New Mexico, USA). email . Cave microbiology and geomicrobiology. Affiliation: New Mexico Institute of Mining and Technology.

  • Bowen-Walker, Dr. P.L. South Wales (UK). email . South Wales (UK) cave fauna

  • Brack, Jr., Dr. Virgil(USA). email . Wintering populations of endangered bats and temperatures of hibernation

  • Brigham, Dr. R. Mark (Saskatchewan, Canada). email . Predator-prey interactions in bats and in nocturnal insectivorous birds. Affiliation: University of Regina.

  • Brown, Dr. Les (North Carolina, USA). email . Cave harvestman-spiders (Opiliones). Affiliation: Gardner-Webb University.

  • Buchanan, Michael (England). email . Cave Ecologist; UK cave dwelling arachnids

  • Camassa, Dott. Michele M. (Italy). email . Director: Museo del Sottosuolo. Ethology and ecology of cave invertebrates, particularly Troglophilus andreinii (Orthoptera, Rhaphidophoridae) Affiliation: Istituto Sperimentale di Biologia del Sottosuolo

  • Chandler, Dr. Donald S.(New Hampshire, USA). email . Entomologist; research has focussed on documenting the insect fauna of old-growth or unique habitats including caves. Cavernicolous beetles. Affiliation: University of New Hampshire.

  • Clarke, Arthur (Tasmania). email . Tasmanian biospeleology. Conservation.

  • Cokendolpher, Mr. James C. (Texas, USA). e-mail . Taxonomy of spiders and harvestmen, invertebrate cave fauna (fossil and recent) of southeastern New Mexico. Affiliation: Midwestern State University. Consulting biologist, Taxonomist and scientific illustrator.

  • Contos, Dr. Annalisa (Australia). email. Biomineralisation in caves. Affiliation: University of Sydney.

  • Crawford, Rod (Washington, USA). email . Invertebrates (especially arachnids) in caves from the North American Pacific Northwest. Affiliation: Burke Museum, University of Washington, Seattle.

  • Culver, Dr. C. David (Washington D.C., USA). email. Adaptive processes in space and time. Affiliation: American University.

  • Dittmar de la Cruz, Dr. Katharina (Laramie, Wyoming, USA). email. Affiliation: University of Wyoming.

  • Elliott, Dr. William (Missouri,USA). email . Conservation and documentation of the cave fauna of Missouri. Affiliation: Missouri Dept. of Conservation.

  • Espinasa, Luis (México City, México). email . cave fish, Apterygota, marine worms from caves Affiliation: Universidad Autonoma del Estado de Morelos

  • Febbroriello, Peter (Connecticut, USA). email . Cave mycology and microbiology.

  • Fenton, Dr. M. Brock (Ontario, Canada). e-mail . Behaviour, ecology and evolution of bats. Affiliation: University of Western Ontario.

  • Ferguson, Lynn (Virginia, USA). email . Cave diplura Affiliation: Longwood College.

  • Helf, Dr. Kurt Lewis (Mammoth Cave, KY). email . cave invertebrate monitoring and cave cricket (Hadenoecus subterraneus) foraging behavior. Affiliation: National Park Service, Mammoth Cave.

  • Hobbs, Dr. H. Horton (Ohio, USA). email . aquatic cave ecosystems; adaptation and evolution of cavernicoles. Affiliation: Wittenberg University.

  • Holler, Dr. Cato O. (Carolina, USA). email . Documentation of the biological resources of North and South Carolina caves. Affiliation: Director, Carolina Cave Survey.

  • Holmberg, Dr. Robert (Alberta, Canada). email . Arachnids. Harvest-spiders in caves. Affiliation: Athabasca University.

  • Holsinger, Dr. John R. (Virginia, USA). email . Subterranean amphipoda. Affiliation: Old Dominion University.

  • Howarth, Dr. Frank G.(Hawaii, USA). email . Cave fauna of Hawaiian lava tubes. Affiliation: Bernice P. Bishop Museum Honolulu.

  • Hubbard, David (Virginia, USA). email . Documentation of the biological resources of Virginia caves. Affiliation: Virginia Natural Heritage Program.

  • Humphreys, Dr. William F. (Bill) (Western Australia). email . Terrestrial invertebrate cave fauna of Western Australia, especially Cape Range. Affiliation: Western Australian Museum.

  • Iliffe, Dr. Thomas (Texas, USA).email . Biology of tropical marine and anchialine caves. Affiliation: Texas A and M University.

  • Jasinska, Dr. Edyta J. (Alberta, Canada). email . Root-driven cave stream ecosystems; Cave and Basin ecosystem in Banff National Park. Affiliation: Golder Associates Ltd.

  • Kingsley, Dr. Kenneth J. (Arizona and Texas, USA). email . Conservation of cave fauna. Preserve design. Bats, especially their use of inactive mines (artificial caves). Affiliation: SWCA, Inc. Environmental Consultants.

  • Krejca, Jean (Texas, USA). email . Cave catfish (Prietella), cirolanid isopods, Mexican and Texas cave fauna. Affiliation: University of Texas, Zara Environmental.

  • Larimore, Richard (Illinois, USA). Ecologist; resource management (including caves in Illinois). Affiliation: retired but active in research.

  • Lebreton, Bernard (France). email . Worldwide bibliography (since 1991). Research reprints. Affiliation: member of Societe de Biospeologie.

  • Lewis, Julian (Jerry) (Indiana, USA). email . Cave biological surveys and consulting; cave isopoda Affiliation: J. Lewis and Associates, Biological Consulting.

  • Longley, Glen (Texas, USA). email . Director of the Edwards Aquifer Research Center at Southwest Texas State University in San Marcos; involved in study and protection of fauna of the Edwards Aquifer.

  • Manolache, Elena (Romania). email . Cave microbiology.

  • Margus, Dr.Mirian D. (Brazil). email . Temporal approaches to adaptive processes. Affiliation: Museu de Zoologia du Universidade de Sao Paulo.

  • Moseley, Max (Songkhla, Thailand). email. Interests: Distribution and ecology of cave invertebrates in Atlantic Canada, NW England and peninsula Thailand/Malaysia, especially in seacaves and guano. Affiliation: Nova Scotia Museum of Natural History.

  • Moulds, Tim (Australia). email. Guano-associated cave invertebrates Affiliation: The University of Adelaide, Adelaide

  • Mugue, Dr. Nikolai S (Russia). email . Affiliation: Russian Academy of Science.

  • Myburgh, Jan (South Africa). email . Cave conservation, ecology and biospeleology.

  • Nagorsen, Dr. David (British Columbia, Canada). email . Systematics, biogeography and conservation of mammals - including bats - of the Pacific Northwest. Affiliation: Mammalia Consulting, Victoria, BC.

  • Pearson, Dr. William A (Kentucky, USA). email . Cave fishes. Affiliation: Clay Fish Collection.

  • Pesce, Giuseppe(Italy).email . systematics and biogeography of crustaceans (copepods, asellid and microparasellid isopods, amphipods of the genera Hadzia and Bogidiella, mysids). Biospeleology. Ground water biology. Affiliation: University of L'Aquila.

  • Poulson, Dr. Thomas (Illinois, USA). email . Ecology of the Mammoth Cave system, Kentucky.

  • Price, Liz (Kuala Lumpur, Malaysia). email . Malaysian biospeleology.

  • Proudlove, Graham Steven (England). email . Cavernicolous fishes. Affiliation: University of Manchester Institute of Science and Technology.

  • Reddell, James R. (Texas, USA). email . Distribution of Texas and Mexico cave fauna and systematics of Schizomida (Arachnida). Affiliation: Texas Memorial Museum, University of Texas at Austin.

  • Richards, Ronald T. (San Juan, Puerto Rico). email . Interests in the life in Puerto Rican caves and in studying non-flying mammals found in the complete darkness of caves worldwide. Affiliation: Sociedad Espeleológica de Puerto Rico, P.O. Box 31074 San Juan, PR 00929.

  • Romero, Dr. Aldemaro (Arkansas, USA). email. Cave Fishes. Affiliation: Arkansas State University

  • Schwendinger, Dr Peter J (Austria). email . Spiders. incl. Liphistus. Affiliation: Innsbruck University.

  • Sbordoni, Professor Valerio (Italy). email . Evolutionary biology of cave animals. Affiliation: University of Roma "Tor Vergata."

  • Shaw, Patrick (British Columbia, Canada). email . Aquatic crustacea. Western Canada and elsewhere, biological inventory/survey.

  • Sket, Boris (Slovenia). email . - biogeography of karst regions, particularly Dinaric karst; origins and ecology of cave faunas; anchihaline caves; taxonomy of Isopoda aquatica, Amphipoda, Hirudinea. Affiliation: Department of Biology, University of Ljubljana Ljubljana,SLOVENIA

  • Dave Slaney (Australia). email . Molecular and morphological evolution and adaptation of cave invertebrates. Affiliation: Dept of Zoology at James Cook University Queensland

  • Steiner, Helmut (Germany). email . Liphistus(Trapdoor Spiders) of South East Asia. Affiliation: Dept. Zoology, Frankfurt University.

  • Stoch, Dr. Fabio (Italy). email . Taxonomy of copepods. Cladistic biogeography and distributional ecology of groundwater taxa in southern Europe. Affiliation: research associate Natural History Museum in Trieste.

  • Taylor, Steve (Illinois, USA). email . Cave fauna of North America. Ecology and biology of Heteroptera. Affiliation: Center for Biodiversity, Illinois Natural History Survey.

  • Thurgate, Mia (Australia). email . Cave ecology and aquatic cave communities, Australia. Affiliation: James Cook University.

  • Trajano, Eleonora (Brazil). email .

  • Wagner, Dr. John A. (Illinois). email . Terrestrial cave communities. Coleoptera. Affiliation: Field Museum of Natural History.

  • Zaragoza, Juan A. (Alicante, Spain). email . Cave-dwelling pseudoscorpions. Affiliation: Departamento de Ecología, Universidad de Alicante, Spain.

This page maintained by Patrick Shaw, Vancouver, BC. Last Updated .